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Elulookirjeldus (CV)
1.Eesnimi Peeter
2.Perekonnanimi Hõrak
3.Töökoht Tartu Ülikool, Zooloogia ja Hüdrobioloogia Instituut, loomaökoloogia õppetool
4.Ametikoht vanemteadur
5.Sünniaeg 11.12.1963 (päev.kuu.aasta)
6.Haridus Dipl. bioloog-zooloog, Tartu Ülikool 1989
7.Teenistuskäik Nõukogude Armee 1983-1985; TA ZBI Ornitoloogia Laboratoorium, insener: sept. 1989 - dets. 1989; TA ZBI Ornitoloogia Laboratoorium, aspirant: dets. 1989 - sept. 1991; TÜ ZHI Loomaökoloogia Õppetool, doktorant: sept. 1991 - sept. 1995; Turku University, Laboratory of Ecological Zoology: 1993 jaan. - dets. (Kone Foundation'i stipendiaat); ZBI Loomaökoloogia Osakond, teadur: jaan. 1996 - märts 1998; Uppsala University, Department of Zoology: külalisteadlane veebr. - märts 1996, veebr.- märts 1997, okt. - nov. 1998; EPMÜ ZBI Zooloogia Osakond, vanemteadur: aprill 1998 - juuli 1999. TÜ ZHI Loomaökoloogia Õppetool, vanemteadur: alates juuli 1999.
8.Teaduskraad PhD
9.Teaduskraadi välja
andnud asutus, aasta
Tartu Ülikool, 1995
10.Tunnustused EV Teaduspreemia 2002 "Baasuuringud evolutsioonilise loomaökoloogia vallas - bioloogilise mitmekesisuse huvides" (koos Raivo Männi; Indrek Otsa ja Toomas Tammaruga)
11.Teadusorganisatsiooniline
ja –administratiivne
tegevus
Rahvusvaheliste teadusajakirjade toimetuskollegiumite liige:

Avian Science (2001-2003)
Ardeola (alates 2003)
Functional Ecology (alates 2005)

Teadusliku programmi komitee liige: 7th congress of the European Ornithologists' Union, Austria 2007

Retsenseerinud käsikirju ajakirjadele Proceedings of the Royal Society of London. B, Biology Letters, Ecology Letters, Naturwissenschaften, Evolution, Journal of Animal Ecology, Functional Ecology, Oecologia, Biological Journal of Linnean Society, Ethology, Ethology Ecology & Evolution, Behavioural Ecology and Sociobiology, Physiology & Behaviour, Ecography, Chemoecology, Polar Biology, Physiological and Biochemical Zoology, Canadian Journal of Zoology, Annales Zoologici Fennici, Journal of Avian Biology, Condor, Avian Science, Ardea, Ardeola, Ornis Fennica, Acta Ornithologica, Journal of Field Ornithology.

Retsenseerinud grantitaotlusi Eesti Teadusfondile

Evalueerinud doktoritööd: Markus Rantala “Immunocompetence and sexual selection in insects” (University of Jyväskylä, 2002).

Oponeerinud doktoritööd: Paola Ninni "Function of carotenoids in barn swallow Hirundo rustica" (Universite Paris 6, 2003).

TÜ Bio-Geo teaduskonna nõukogu liige (alates 2003) ja ETF Bio-Geoteaduste ekspertkomisjoni liige (alates 2003)
12.Juhendamisel kaitstud
väitekirjad

Lauri Saks, PhD, 2004, juh. Peeter Hõrak. Immune function, parasites, and carotenoid-based ornaments in greenfinches. Tartu

Ulvi Karu, MSc, 2004, juh. Peeter Hõrak. Eksperimentaalse koktsiidinakkuse mõju rohevintide (Carduelis chloris) tervislikule seisundile ja sulestiku värvusele. Tartu

Helen Vellau, MSc, 2002, juh. Peeter Hõrak. Karotinoididel baseeruv sulestiku värvus ja maternaalsed mõjud rasvatihasel. Tartu

Lauri Saks, MSc, 2002, juh. Peeter Hõrak. Immuunparameetrid ja muud konditsiooni-indeksid vangistuses peetavatel rohevintidel: püsivus ajas, seos sulestiku värvusega ja reaktsioon humoraalsele immuunväljakutsele. Tartu

Lea Tegelmann, MSc, 2001, juh. Peeter Hõrak. Leukotsüütsed immunparameetrid rasvatihase pesapoegadel: individuaalse varieeruvuse põhjused ja tagajärjed. Tartu

Indrek Ots, PhD, 1999, juh. Peeter Hõrak. Health state indices of reproducing great tits (Parus major): sources of variation and connections with life-history traits. Tartu

Indrek Ots, MSc, 1997, juh. Peeter Hõrak. Hematological health state indices of reproducing great tits. Sources of natural variation and a response to brood size manipulation. Tartu

13.Teadustöö põhisuunad Vanemlike investeeringute teooria: Sigimise hinna maksmise käitumisökoloogilised aspektid ja füsioloogilised mehhanismid. Immuunökoloogia: Immunvastuse hind. Immuunfunktsioon, parasiidiresistentsus ja karotinoididel põhinevad signaalid sugulise ning loodusliku valiku kontekstis. Kliinilised vereparameetrid isendi konditsiooni ja parasiidinakkuse indikaatoritena loomaökoloogilistes mudelsüsteemides. Vereparasiitide, koktsiidide ja nematoodide nakkuse mõju peremeeste kohasusele ja füsioloogiale. Oksüdatiivne stress ökoloogilises kontekstis.
14.Jooksvad grandid Immuunfunktsioon ökoloogilises kontekstis: oksüdatiivne stress, karotinoidsed signaalid ja parasiidiresistentsus (ETF grant # 6222; 2005-2008)
15.Teaduspublikatsioonid

Hõrak, P., Tummeleht, L. & Talvik, H. (in press) Predator threat, copulation effort, and immunity in male rats. Journal of Zoology

Hõrak, P., Tummeleht, L. & Talvik, H. (in press) Predictors and markers of resistance to neurotropic nematode infection in rodent host. Parasitology Research

Saks, L., Karu, U., Ots, I., Hõrak, P. (2006) Do standard measures of immunocompetence reflect parasite resistance? The case of greenfinch coccidiosis. Functional Ecology (in press)

Hõrak P., Saks L., Karu U. & Ots I. (2005) Host resistance and parasite virulence in greenfinch coccidiosis. Journal of Evolutionary Biology in press (doi:10.1111/j.1420-9101.2005.00988.x)

Hõrak P., Saks L., Karu U., Ots I., Surai P. F., & McGraw, K. J. (2004) How coccidian parasites affect health and appearance of greenfinches. Journal of Animal Ecology 73, 935-937

Hõrak, P., Surai, P. F., Ots, I. & Møller, A. P. (2004) Fat soluble antioxidants in brood-rearing great tits: relations to health and appearance. Journal of Avian Biology 35, 63-70.

Freitak, D., Ots, I., Vanatoa, A. & Hõrak, P. (2003) Immune response is energetically costly in white cabbage butterfly pupae. Proceedings of the Royal Society of London Series B Supplementum (Biology Letters) 270, S220-222.

Hõrak, P. (2003) When to pay the cost of reproduction? A brood size manipulation experiment in great tits (Parus major). Behavioral Ecology and Sociobiology 54, 105-112.

Hõrak, P. & Saks, L. (2003) Animal allure and health linked by plant pigments. BioEssays 25, 746-747.

Hõrak, P., Saks, L., Ots, I., Kullisaar, T., Kollist, H. & Zilmer, M. (2003) Physiological effects of immune challenge in captive greenfinches (Carduelis chloris). Canadian Journal of Zoology 81, 371-379.

Saks, L., McGraw, K. J. M. & Hõrak, P. (2003) How feather colour reflects its carotenoid content. Functional Ecology 17, 555-561.

Saks, L., Ots, I., Hõrak, P. (2003) Carotenoid-based plumage coloration of male greenfinches reflects health and immunocompetence. Oecologia 134, 301-307.

Hõrak, P., Saks, L., Ots, I. & Kollist, H. (2002) Repeatability of condition indices in captive greenfinches (Carduelis chloris). Canadian Journal of Zoology 80, 636-643.

Hõrak, P., Surai, P. F. & Møller, A. P. (2002) Fat-soluble antioxidants in the eggs of great tits Parus major in relation to breeding habitat and laying sequence. Avian Science 2, 123-130

Hõrak, P., Ots, I., Vellau, H., Spottiswoode, C. & Møller, A. P. (2001) Carotenoid-based plumage coloration reflects hemoparasite infection and local survival in breeding great tits. Oecologia 123, 166-173.

Ots, I., Kerimov, A. B, Ivankina, E. V., Ilyina, T. A., Hõrak, P. (2001) Immune challenge affects basal metabolic activity in wintering great tits. Proceedings of the Royal Society of London Series B 268, 1475-1482.

Hõrak, P., Ots, I., Tegelmann, L. & Møller, A.P. (2000) Health impact of phytohaemagglutinin-induced immune challenge on great tit nestlings. Canadian Journal of Zoology 78, 905-910.

Hõrak, P.,Vellau, H., Ots, I., & Møller, A. P. (2000) Growth conditions affect carotenoid-based plumage coloration of great tit nestlings Naturwissenschaften 87, 460-464.

Hõrak, P., Jenni-Eiermann, S., & Ots, I. (1999) Do great tits (Parus major) starve to reproduce? Oecologia 119, 293-299.

Hõrak, P., Tegelmann, L., Ots, I., & Møller, A.P. (1999) Immune function and survival of great tit nestlings in relation to growth conditions. Oecologia 121, 316-322.

Tammaru, T. & Hõrak, P. (1999) Should one invest more in large broods? Not necessarily. Oikos 85, 574-81.

Hõrak, P., Jenni-Eiermann, S., Ots, I., & Tegelmann, L. (1998) Health and reproduction: sex-specific clinical profile of Great Tits (Parus major) in relation to breeding. Canadian Journal of Zoology 76, 2235-2244.

Hõrak, P. & Lebreton, J.-D. (1998) Survival of adult Great Tits in relation to sex and habitat: a comparison of urban and rural populations. Ibis140, 205-209.

Hõrak, P. & Ots, I. (1998) Health state and local survival in the Great Tit. Biol. Cons. Fauna 102, 307-311.

Hõrak, P., Ots, I., & Murumägi, A. (1998) Hematological health state indices of reproducing Great Tits: a response to brood size manipulation. Functional Ecology 12 , 750-756.

Ots, I. & Hõrak, P. (1998) Health impact of blood parasites in breeding great tits. Oecologia 116, 441-448.

Ots, I., Murumägi, A., & Hõrak, P. (1998) Hematological health state indices of reproducing Great Tits: methodology and sources of natural variation. Functional Ecology 12, 700-707.

Hõrak, P., Mänd, R., & Ots, I. (1997) Identifying targets of selection: a multivariate analysis of reproductive traits in the Great Tit. Oikos 78, 592-600.

Hõrak, P. & Tammaru, T. (1996) Between-year variation in breeding conditions biases heritability estimates for body size in birds. Ardea 84, 127-135.

Ots, I. & Hõrak, P. (1996) Great Tits Parus major trade health for reproduction. Proceedings of the Royal Society of London Series B 263, 1443-1447.

Hõrak, P. (1995) Brood reduction facilitates female but not offspring survival in the great tit. Oecologia 102, 515-519.

Hõrak, P., Mänd, R., Ots, I., & Leivits, A. (1995) Egg size variation in the Great Tit: individual, habitat and geographic differences. Ornis Fennica 73, 97-114.

Hõrak, P. (1994) Effect of nestling history on adult size and reproduction in the Great Tit. Ornis Fennica 71, 47-54.

Hõrak, P. (1993) Low fledging success of urban Great Tits. Ornis Fennica 70, 168-172.

viimati muudetud: 26.09.2005

Curriculum Vitae (CV)
1.First Name Peeter
2.Surname Hõrak
3.Institution Tartu University, Institute of Zoology and Hydrobiology, Chair of Animal Ecology
4.Position senior researcher
5.Date of birth 11.12.1963 (day.month.year)
6.Education diploma degree in biology (zoology)in Tartu University
7.Research and
professional experience
Military service 1983-1985; Inst. of Zool. and Botany (ZBI) Lab. of Ornithology, lab. assistant, Sept. 1989 - Dec. 1989; ZBI Lab. of Ornithology, postgrad. student Dec. 1989 - Sept. 1991; Tartu University, Ph.D. studies in animal ecology, Sept. 1991 - Sept. 1995; ZBI Animal Ecology Section, researcher, Jan. 1996 - March 1998; ZBI Zoology section, senior researcher, Apr. 1998 - June 1999; Tartu University, Inst. of Zool. and Hydrobiol., senior researcher, July 1999 -
Research training: 1991 Dec., Course in biostatistics in Reposaari (Turku University); 1993 Jan. - Dec., Turku University, Lab. of Ecological Zoology; 1996 Feb.- March, 1997 Feb.- March; 1998 October - November Uppsala Univ., Department of Zoology
8.Academic degree PhD
9.Dates and sites of
earning the degrees
Tartu University, 1995
10.Honours/awards Annual Science Award of Estonian Republic in Biological and Geosciences in 2002 (together with Raivo Mänd, Indrek Ots and Toomas Tammaru)
11.Research-administrative
experience
associate editor:

Avian Science 2001-2003
Ardeola since 2003
Functional Ecology since 2005

Member of the scientific programme committee: 7th congress of the European Ornithologists' Union, Austria 2007

Reviewer: Proceedings of the Royal Society of London. B, Biology Letters, Ecology Letters, Naturwissenschaften, Evolution, Journal of Animal Ecology, Functional Ecology, Oecologia, Biological Journal of Linnean Society, Ethology, Ethology Ecology & Evolution, Behavioural Ecology and Sociobiology, Physiology & Behaviour, Ecography, Chemoecology, Polar Biology, Physiological and Biochemical Zoology, Canadian Journal of Zoology, Annales Zoologici Fennici, Journal of Avian Biology, Condor, Avian Science, Ardea, Ardeola, Ornis Fennica, Acta Ornithologica, Journal of Field Ornithology.

Reviewed grant applications for Estonian Science Foundation

Reviewed Markus Rantala's PhD thesis “Immunocompetence and sexual selection in insects” (University of Jyväskylä, 2002).

Opponent to Ph.D. theses of Paola Ninni “The function of carotenoids in the barn swallow (Hirundo rustica)” ( Universite Paris6, 2003).

Member of the Council of Estonian Science Foundation (Bio- and Geosciences, since 2003) and member of the Council of Faculty of Biology and Geography, Tartu University (since 2003)
12.Supervised dissertations

Lauri Saks, PhD, 2004, superv. Peeter Hõrak. Immune function, parasites, and carotenoid-based ornaments in greenfinches. Tartu

Ulvi Karu, MSc, 2004, superv. Peeter Hõrak. Eksperimentaalse koktsiidinakkuse mõju rohevintide (Carduelis chloris) tervislikule seisundile ja sulestiku värvusele. Tartu

Helen Vellau, MSc, 2002, superv. Peeter Hõrak. Karotinoididel baseeruv sulestiku värvus ja maternaalsed mõjud rasvatihasel. Tartu

Lauri Saks, MSc, 2002, superv. Peeter Hõrak. Immuunparameetrid ja muud konditsiooni-indeksid vangistuses peetavatel rohevintidel: püsivus ajas, seos sulestiku värvusega ja reaktsioon humoraalsele immuunväljakutsele. Tartu

Lea Tegelmann, MSc, 2001, superv. Peeter Hõrak. Leukotsüütsed immunparameetrid rasvatihase pesapoegadel: individuaalse varieeruvuse põhjused ja tagajärjed. Tartu

Indrek Ots, PhD, 1999, superv. Peeter Hõrak. Health state indices of reproducing great tits (Parus major): sources of variation and connections with life-history traits. Tartu

Indrek Ots, MSc, 1997, superv. Peeter Hõrak. Hematological health state indices of reproducing great tits. Sources of natural variation and a response to brood size manipulation. Tartu

13.Current research program Parental Investment theory: mechanisms for costing reproduction. Costs of immune responses. Immunoecology of parasite resistance and carotenoid-based signals. Applicability of clinical health state indices in ecological research. Physiological and fitness-related impacts of blood parasites, coccidians and nematodes on hosts. Oxidative stress in ecological context.
14.Current grant funding Immune function in the ecological context: oxidative stress, carotenoid-based signals and parasite resistance (ESF grant # 6222; 2005-2008)
15.List of most important publications

Hõrak, P., Tummeleht, L. & Talvik, H. (in press) Predator threat, copulation effort, and immunity in male rats. Journal of Zoology

Hõrak, P., Tummeleht, L. & Talvik, H. (in press) Predictors and markers of resistance to neurotropic nematode infection in rodent host. Parasitology Research

Saks, L., Karu, U., Ots, I., Hõrak, P. (2006) Do standard measures of immunocompetence reflect parasite resistance? The case of greenfinch coccidiosis. Functional Ecology (in press)

Hõrak P., Saks L., Karu U. & Ots I. (2005) Host resistance and parasite virulence in greenfinch coccidiosis. Journal of Evolutionary Biology in press (doi:10.1111/j.1420-9101.2005.00988.x)

Hõrak P., Saks L., Karu U., Ots I., Surai P. F., & McGraw, K. J. (2004) How coccidian parasites affect health and appearance of greenfinches. Journal of Animal Ecology 73, 935-937

Hõrak, P., Surai, P. F., Ots, I. & Møller, A. P. (2004) Fat soluble antioxidants in brood-rearing great tits: relations to health and appearance. Journal of Avian Biology 35, 63-70.

Freitak, D., Ots, I., Vanatoa, A. & Hõrak, P. (2003) Immune response is energetically costly in white cabbage butterfly pupae. Proceedings of the Royal Society of London Series B Supplementum (Biology Letters) 270, S220-222.

Hõrak, P. (2003) When to pay the cost of reproduction? A brood size manipulation experiment in great tits (Parus major). Behavioral Ecology and Sociobiology 54, 105-112.

Hõrak, P. & Saks, L. (2003) Animal allure and health linked by plant pigments. BioEssays 25, 746-747.

Hõrak, P., Saks, L., Ots, I., Kullisaar, T., Kollist, H. & Zilmer, M. (2003) Physiological effects of immune challenge in captive greenfinches (Carduelis chloris). Canadian Journal of Zoology 81, 371-379.

Saks, L., McGraw, K. J. M. & Hõrak, P. (2003) How feather colour reflects its carotenoid content. Functional Ecology 17, 555-561.

Saks, L., Ots, I., Hõrak, P. (2003) Carotenoid-based plumage coloration of male greenfinches reflects health and immunocompetence. Oecologia 134, 301-307.

Hõrak, P., Saks, L., Ots, I. & Kollist, H. (2002) Repeatability of condition indices in captive greenfinches (Carduelis chloris). Canadian Journal of Zoology 80, 636-643.

Hõrak, P., Surai, P. F. & Møller, A. P. (2002) Fat-soluble antioxidants in the eggs of great tits Parus major in relation to breeding habitat and laying sequence. Avian Science 2, 123-130

Hõrak, P., Ots, I., Vellau, H., Spottiswoode, C. & Møller, A. P. (2001) Carotenoid-based plumage coloration reflects hemoparasite infection and local survival in breeding great tits. Oecologia 123, 166-173.

Ots, I., Kerimov, A. B, Ivankina, E. V., Ilyina, T. A., Hõrak, P. (2001) Immune challenge affects basal metabolic activity in wintering great tits. Proceedings of the Royal Society of London Series B 268, 1475-1482.

Hõrak, P., Ots, I., Tegelmann, L. & Møller, A.P. (2000) Health impact of phytohaemagglutinin-induced immune challenge on great tit nestlings. Canadian Journal of Zoology 78, 905-910.

Hõrak, P.,Vellau, H., Ots, I., & Møller, A. P. (2000) Growth conditions affect carotenoid-based plumage coloration of great tit nestlings Naturwissenschaften 87, 460-464.

Hõrak, P., Jenni-Eiermann, S., & Ots, I. (1999) Do great tits (Parus major) starve to reproduce? Oecologia 119, 293-299.

Hõrak, P., Tegelmann, L., Ots, I., & Møller, A.P. (1999) Immune function and survival of great tit nestlings in relation to growth conditions. Oecologia 121, 316-322.

Tammaru, T. & Hõrak, P. (1999) Should one invest more in large broods? Not necessarily. Oikos 85, 574-81.

Hõrak, P., Jenni-Eiermann, S., Ots, I., & Tegelmann, L. (1998) Health and reproduction: sex-specific clinical profile of Great Tits (Parus major) in relation to breeding. Canadian Journal of Zoology 76, 2235-2244.

Hõrak, P. & Lebreton, J.-D. (1998) Survival of adult Great Tits in relation to sex and habitat: a comparison of urban and rural populations. Ibis140, 205-209.

Hõrak, P. & Ots, I. (1998) Health state and local survival in the Great Tit. Biol. Cons. Fauna 102, 307-311.

Hõrak, P., Ots, I., & Murumägi, A. (1998) Hematological health state indices of reproducing Great Tits: a response to brood size manipulation. Functional Ecology 12 , 750-756.

Ots, I. & Hõrak, P. (1998) Health impact of blood parasites in breeding great tits. Oecologia 116, 441-448.

Ots, I., Murumägi, A., & Hõrak, P. (1998) Hematological health state indices of reproducing Great Tits: methodology and sources of natural variation. Functional Ecology 12, 700-707.

Hõrak, P., Mänd, R., & Ots, I. (1997) Identifying targets of selection: a multivariate analysis of reproductive traits in the Great Tit. Oikos 78, 592-600.

Hõrak, P. & Tammaru, T. (1996) Between-year variation in breeding conditions biases heritability estimates for body size in birds. Ardea 84, 127-135.

Ots, I. & Hõrak, P. (1996) Great Tits Parus major trade health for reproduction. Proceedings of the Royal Society of London Series B 263, 1443-1447.

Hõrak, P. (1995) Brood reduction facilitates female but not offspring survival in the great tit. Oecologia 102, 515-519.

Hõrak, P., Mänd, R., Ots, I., & Leivits, A. (1995) Egg size variation in the Great Tit: individual, habitat and geographic differences. Ornis Fennica 73, 97-114.

Hõrak, P. (1994) Effect of nestling history on adult size and reproduction in the Great Tit. Ornis Fennica 71, 47-54.

Hõrak, P. (1993) Low fledging success of urban Great Tits. Ornis Fennica 70, 168-172.

last updated: 26.09.2005

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